HOORAY --- A coherent answer. And, given that answer, what determines the difference between the Vbe of those two transistors? Maybe the bias resistors have some effect. Certainly does not seem like a robust design with respect to class AB quiescent final stage current. I consider it a sloppy class B design. Some will work well, some not so well. Depends on Vbe matching of discrete devices.
As for the 220K resistors, I don't can't assign a purpose to those either.
Another quality of design issue is that the opamp is not sourcing current at 0V. I generally add a load resistor from the opamp output to the negative supply to keep the output transistor conducting. Helps with crossover distortion.
The lack of bypass on the feedback resistor has been mentioned before.
I don't have an answer for why lowering the bias resistors to 1.5K made the sound poor. I'd have to see the waveform. These transistors have internal resistors that are not on your schematic. Maybe that is part of it. ( see fig 1 in
http://www.onsemi.com/pub_link/Collateral/TIP120-D.PDF )